A novel cell adhesive basement membrane protein with a putative role in stimulus-secretion coupling by lacrimal acinar exocrine cells. Laurie, Gordon W., J. Douglas Glass, Rebecca A. Ogle, Cameron M. Stone, James R. Sluss, and Lanlin Chen. Department of Cell Biology, University of Virginia, Charlottesville, VA
APStracts 2:0419C, 1995.
Regulated secretion requires the developmental coupling of neuronal or hormonal stimuli to an exocytotic response - a multistep pathway whose appearance may be linked with cellular adhesion to the newly formed exocrine cell basement membrane. We screened for adhesion -associated coupling activity using lacrimal acinar cells and have identified BM180', a novel basement membrane protein enriched in guanidine-HCl extracts of lacrimal and parotid exocrine secretory glands. BM180 resides primarily in a previously unexamined lower molecular weight basement membrane peak (Pk 2') which contains cell adhesion activity inhibitable with the anti-BM180 monoclonal antibody 3E12. Removal of Pk 2 by gel filtration or preincubation of basement membrane with 3E12 decreased regulated peroxidase secretion by one -half without affecting constitutive secretion nor the amount of cellular peroxidase available for release. Adding back Pk 2 restored regulated secretion in a dose dependent and 3E12 inhibitable manner; and suggested a synergistic relationship between BM180 and laminin-1. BM180 has a mobility of 180 kDa and 60 kDa in the absence or presence of DTT, respectively; and shows no immunological identity by competitive ELISA with laminin-1, collagen IV, entactin, fibronectin, BM-40, perlecan or vitronectin. We propose that BM180 is an important resident of certain glandular basement membranes where it interacts with the cell surface thereby possibly signalling the appearance of a transducing element in the stimulus-secretion coupling pathway.

Received 28 September 1995; accepted in final form 13 November
1995.
APS Manuscript Number C594-5.
Article publication pending Am. J. Physiol. (Cell Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on 8 December 95