Deoxygenation-induced cation fluxes in sickle cells iv. modulation by external calcium.. Joiner, Clinton H., Maorong Jiang, and Robert S. Franco. Departments of Pediatrics, Physiology and Biophysics, and Internal Medicine, University of Cincinnati College of Medicine, Children's Hospital Medical Center, Cincinnati Comprehensive Sickle Cell Center, Cincinnati, Ohio
APStracts 2:0095C, 1995.
Net cation movements were measured in low density sickle red blood cells (SS RBC) in the presence and absence of oxygen. External Ca++ (Cao++) partially inhibited deoxygenation-induced fluxes of both Na+ and K+. Deoxygenation-induced Na+ influx was reduced by 2 mM Cao++ to 0.71 +/- 0.04 (SEM) of its value in Ca++-free solutions, whereas this ratio was 0.90 +/- 0.05 for K+ efflux (p<0.01 by paired t-test). Since Cao++ inhibited Na+ influx more than K+ efflux, net cation loss in deoxygenated SS RBC was higher in the presence of Cao++. In separate experiments, Cao++ reduced deoxygenation-induced Na+ influx to 0.66 +/- 0.03 of its Ca++-free value compared to 0.77 +/- 0.03 for Rb+ influx (p< 0.001), indicating relative selectivity of this effect for Na+ over Rb+. However, this effect is not specific for Ca++, as other divalent cations also inhibited deoxygenation-induced Na+ and K+ fluxes. Under the conditions of these experiments, no evidence for K+ channel activation was found, indicating that K+ loss measured in deoxygenated SS RBC was mediated by the deoxygenation-induced pathway. These studies show that in the presence of Cao++, deoxygenation-induced Na+ influx and K+ efflux are unbalanced. This pathway can, therefore, mediate cation loss and contribute directly to cellular dehydration in SS RBC.

Received 17 November 1994; accepted in final form 13 January
1995.
APS Manuscript Number C675-4.
Article publication pending Am. J. Physiol. (Cell Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on 28 February 1995.