Nuclear responses to depletion of mitochondrial dna in human cells. Li, Kang, P. Darrell Neufer, and R. Sanders Williams. Departments of Internal Medicine and Biochemistry, The University of Texas Southwestern Medical Center at Dallas, Dallas, TX 75235 -8573
APStracts 2:0206C, 1995.
The derivation of human cell lines devoid of mtDNA (r0) provides an opportunity to study nuclear responses to a chronic impairment of mitochondrial oxidative phosphorylation. Expression of several nuclear genes is induced in human r0 cells, including those encoding integral proteins of the mitochondrial inner membrane, intermediate filaments, and ribosomes. In contrast to conditions in which mitochondrial respiration is altered acutely, expression of heat shock proteins and immediate early genes is not induced. Mitochondria from r0 cells maintain a transmembrane electrochemical potential and are distributed within the cytoplasm of these cells in a manner indistiguishable from wild type cells. We conclude that a chronic deficiency of mitochondrial oxidative phosphorylation produced by elimination of mtDNA is associated with a different pattern of gene induction than that provoked by other acute or subacute conditions that impair mitochondrial respiration or create energy demands in excess of mitochondrial respiratory capacity.

Received 30 November 1994; accepted in final form 11 May 1995.
APS Manuscript Number C701-4.
Article publication pending Am. J. Physiol. (Cell Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on 26 May 1995.