Sphingolipid actions on sodium and calcium currents of rat ventricular myocytes. Yasui, Kenji, and Philip Palade. Departments of Physiology & Biophysics and of Pharmacology & Toxicology, University of Texas Medical Branch, Galveston, TX 77555 -0641, USA
APStracts 2:0321C, 1995.
Sphingosine, an endogenous phospholipid known to produce significant decreases in myoplasmic calcium transients, was shown to have a pronounced inhibitory effect on inward sodium and L-type calcium currents in rat ventricular myocytes. Sphingosine action was accompanied by a slowing of inactivation of both kinds of current. Sphingosine and sphingosylphosphorylcholine (SPC) both caused depolarizing shifts in the activation curves for the two channels. In tests on calcium currents, sphingosine showed no high affinity for inactivated states nor exhibited any use dependence. The mechanism of the blocking action of sphingosine does not appear to involve effects on bulk surface charge nor, at least for calcium channels, a voltage -dependent block. Instead the results appear most consistent with an effect of sphingosine on channel gating. The shift in the voltage dependence of channel activation by sphingosine and SPC appears likely to be a feature of both the hydrocarbon chains and the net positive charge of these amphiphiles.

Received 3 April 1995; accepted in final form 28 August 1995.
APS Manuscript Number C190-5.
Article publication pending Am. J. Physiol. (Cell Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on 23 September 1995.