Temporal and spatial heterogeneities of intracellular ca2+ signalling - their mechanisms and roles in the physiology of the gastrointestinal secretory cells. Toescu, Emil C. Dept. of Physiology, School of Medicine,
APStracts 2:0092G, 1995.
In the cells of the gastrointestinal tract, as in most other types of cells, many of the fundamental cellular functions are mediated by increases of intracellular calcium. These increases are mediated primarily by elevations of the inositol 1,4,5-trisphosphate (InsP3) which, in turn, are a consequence of the activation of specific plasma membrane receptors. Extensive investigations of these intracellular processes in the last few years revealed the complexities of Ca2+ signalling. The extreme heterogeneity of Ca2+ signalling, which is manifest by variations both in the time-course and the spatial distribution of these signals, forms the central theme of this review. On the basis of the available information the various intracellular mechanisms which allow the manifestation of this heterogeneity are discussed. It is argued that these variations in Ca2+ response are the result not only of the existence of multiple, complex mechanisms participating in the generation and maintenance of the agonist-evoked Ca2+ signal but also of the existence of various isoforms for almost each of the main molecules involved in signalling and the concomitant participation of many intracellular modulators.

Received 25 April 1995; accepted in final form 10 May 1995.
APS Manuscript Number G196-5.
Article publication pending Am. J. Physiol. (Gastrointest. Liver
Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on 16 May 1995.