Sarcoplasmic reticulum in cardiac length-dependent activation in rabbits. Bluhm, Wolfgang F., and Wilbur Y. W. Lew. Departments of Bioengineering and Medicine (Cardiology), The University of California, San Diego, La Jolla, California 92093-0412 and Cardiology Division, Department of Veterans Affairs Medical Center, San Diego, California 92161
APStracts 2:0106H, 1995.
After a step increase in length of rabbit right ventricular papillary muscles, active stress increased immediately followed by a further slow increase in stress over 15 to 20 minutes. We studied the contribution of the sarcoplasmic reticulum (SR) to the slow change in stress (SCS) after changing muscle length from 85 % to 95 % of Lmax. The SCS amounted to 32.5 +/- 12.2 % (mean +/- SD, n=19) of the total increase in active stress. This was associated with a 13.2 +/- 8.7 % increase in calcium content of the SR as estimated with rapid cooling contractures (p<0.0001, n=19). However, SCS was not dependent on SR calcium content. There was no significant attenuation in SCS after SR calcium depletion with ryanodine (n=6), SR Ca-ATPase inhibition with cyclopiazonic acid (n=6), or combined treatment with ryanodine and cyclopiazonic acid (n=3). We conclude that in the rabbit, SR calcium content increases slowly following a step increase in cardiac muscle length, but the slow changes in active stress are not dependent on the sarcoplasmic reticulum.

Received 11 August 1994; accepted in final form 17 March 1995.
APS Manuscript Number H716-4.
Article publication pending Am. J. Physiol. (Heart Circ. Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on  4 April 1995.