Oxygen saturation and ph changes in cremaster microvessels of the rat. Kobayashi, Hirosuke, and Naosada Takizawa. Department of Medicine, and Physiology, Kitasato University School of Medicine, Kitasato 1-15-1, Sagamihara, Kanagawa 228, Japan
APStracts 2:0460H, 1995.
Recent studies have shown a considerable extent of diffusive oxygen transfer from pre-capillary vessels, but the occurrence of oxygen influx into post-capillary vessels (diffusive shunting) is still controversial. In this study we investigated diffusive shunting between pre- and post-capillary vessels in muscle tissue and pH changes in the microvessels in an attempt to determine the physiological significance of the parallel arrangement of arterial and venous vessels in tissue oxygenation. We used rat cremaster muscles, and controlled the breathing of anesthetized rats using urethane with FIO2 set at 0.3 and 0.12. We employed a non-invasive spectrophotometric method of measuring pH, using a pH sensitive dye (1-hydroxypyrene-3,6,8-trisulfonic acid) and the oxygen saturation level of hemoglobin (SO2) in microvessels from small arteries down to arterioles, and from venules up to small veins. At FIO2 0.30 pH significantly decreased from 7.39 0.02 (mean S.D.) to 7.26 0.05 in pre-capillary vessels and remained close to constant during passage through post-capillary vessels. At FIO2 0.12 pH decreased from 7.36 0.03 to 7.01 0.06 in pre-capillary vessels and then increased in post-capillary vessels from 6.75 0.19 to 7.27 0.08. At FIO2 0.30 SO2 decreased from 98.6 5.4 to 64.2 4.5% in pre-capillary vessels and increased from 62.5 6.5 to 89.4 4.1% in post-capillary vessels. At FIO2 0.12 SO2 decreased from 47.0 5.5 to 25.1 3.8% in pre-capillary vessels and remained close to constant in post-capillary vessels. In conclusion, during hyperoxia diffusive shunting of oxygen was enhanced, and during hypoxia acid was accumulated in peripheral vessels. It is speculated that acid accumulated due to back-diffusion of CO2 from venous to arterial microvessels, liberating oxygen bound to hemoglobin via the Bohr effect. The diffusive shunting of oxygen and counter-diffusion and accumulation of CO2 may contribute to the homeostasis of tissue oxygenation levels.

Received 5 June 1995; accepted in final form 26 September 1995.
APS Manuscript Number H514-5.
Article publication pending Am. J. Physiol. (Heart Circ. Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on 6 November 95