THE STRUCTURAL BASIS OF CORTICAL SYNCHRONIZATION. II. EFFECTS OF CORTICAL LESIONS. MUNK, M.H.J., NOWAK, L.G., NELSON, J.I., BULLIER J. Cerveau et Vision, INSERM Unit[acute]e 371, 18 avenue du Doyen L[acute]epine, 69500 Bron/Lyon France, t[acute]el: (33) 72 13 15 83, fax: (33) 72 13 15 99.
APStracts 2:0260N, 1995.
SUMMARY AND CONCLUSIONS
1. In order to understand the structural basis of the different types of interhemispheric synchronizations described in the preceding paper, we made sections of the corpus callosum and lesions of extrastriate cortex. We measured the effects of such operations on the frequency of encounter, width and strength of T, C and H peaks in cross-correlation histograms (CCHs) computed from single and multi-unit recordings from areas 17-18 of opposite cortical hemispheres in the cat. 2. Section of the corpus callosum led to a complete abolition of T and C couplings and a strong reduction of encounter rate and strength of H coupling. A section limited to the posterior half of the corpus callosum abolished completely T and C couplings. This suggests that T and C couplings are mediated by the direct reciprocal connections between visual cortical areas circulating through the posterior part of the corpus callosum. 3. The encounter rate of H peaks depended upon the extent of the callosal cut. Larger lesions gave a more pronounced reduction of the number of H peaks. From this observation we conclude that H peaks are, at least partially, mediated by polysynaptic pathways involving widely distributed cortical regions. 4. Extensive lesions of extrastriate cortex were made by aspiration of the grey matter or injections of ibotenic acid. These lesions removed the direct inputs from cortical areas sending ipsilateral as well as contralateral inputs to the area 17-18 border region. Encounter rate and coupling strength of C and H peaks were decreased whereas little effect was observed on T peaks. 5. These results demonstrate that all types of interhemispheric synchronization are mediated by corticocortical connections and that T and C peaks are generated by reciprocal connections between areas 17/18 of each hemisphere. Feedback connections play a role in mediating or facilitating the C and H types of interhemispheric synchronization.

Received 12 May 1995; accepted in final form 27 July 1995.
APS Manuscript Number J258-4.
Article publication pending J. Neurophysiol.
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on 15 September 1995.