Potential role of nadh oxidoreductase-derived reactive o2 species in calf pulmonary arterial po2-elicited responses. Mohazzab-H., Kamal M., Raisa P. Fayngersh, Pawel M. Kaminski, and Michael S. Wolin. Department of Physiology, New York Medical College, Valhalla, NY. 10595
APStracts 2:0104L, 1995.
Our laboratory has previously reported evidence that tone responses of isolated endothelium-removed calf pulmonary arteries elicited by changes in PO2 appear to be mediated via changes in H2O2 and cGMP, and that the PO2 sensor mechanism is hypothesized to involve changes in superoxide anion (O2.-) production by a microsomal NADH -oxidoreductase, which is the major source of O2.- in this tissue detected by lucigenin-elicited chemiluminescence (CL). In this study we examined if the flavoprotein-directed inhibitor of O2.- producing NAD(P)H oxidoreductases, diphenyliodonium (DPI), could be employed as an inhibitor of O2.- production by NADH oxidoreductase which functions as a selective probe for PO2-elicited tone responses in calf pulmonary arterial smooth muscle. It was found that 1 [mu]M DPI inhibited NADH-dependent production of CL in the arterial smooth muscle homogenate by 49% (n=10). DPI reduced basal CL from endothelium-removed pulmonary arteries by 41% (n=15). In endothelium -removed pulmonary arteries precontracted with U46619, the hypoxic contraction of 2.3+/-0.5 g was reduced to 0.1+/-0.4 g (n=7) by DPI and the reoxygenation relaxation of 32.7+/-7.5% was decreased to 4.4+/-1.4% (n=7). DPI did not have any significant effect on U46619 or K- elicited tone generation. DPI also did not alter the relaxation to H2O2 (1 [mu]M-0.1 mM n=6), nitric oxide (0.42 nM-420 nM, n=12) or isoproterenol (1 nM-1 [mu]M, n=6). Probes for other flavoprotein O2.- producing systems, including xanthine oxidase (0.1 mM oxypurinol), nitric oxide synthase (0.1 mM nitro-L-arginine) or mitochondrial electron transport (50 [mu]M rotenone) did not alter the tone response to changes in PO2. These results are consistent with a hypothesized role for H2O2 formation via NADH oxidoreductase-derived O2.- in pulmonary artery responses to changes in PO2.

Received 19 August 1994; accepted in final form 13 June 1995.
APS Manuscript Number L242-4.
Article publication pending Am. J. Physiol. (Lung Cell. Mol.
Physiology).
ISSN 1080-4757 Copyright 1995 The American Physiological Society.
Published in APStracts on  6 July 1995.