Development and initial application of an in vitro model of apoptosis in rodent cholangiocytes. Quea, Florencia G., Gregory J. Goresb, Nicholas F. Larussob. Center for Basic Research in Digestive Diseases, Division of Gastroenterologic and General Surgery, Division of Gastroenterology and Internal Medicine
APStracts 3:0151G, 1996.
While histologic data suggest that cholangiocytes die by apoptosis in human liver diseases, no information exists on the mechanisms of cholangiocyte apoptosis. Thus, our aims were to establish an in vitro model of cholangiocyte apoptosis and to test the hypothesis that changes in intracellular ions would cause apoptosis in cholangiocytes by a protease-sensitive pathway. A large number of pro-apoptotic agents were ineffective in inducing apoptosis in rat or human cholangiocytes in culture; in contrast, beauvericin, a K+ ionophore, caused apoptosis in both cell lines despite their expression of Bcl -2. Although beauvericin decreased intracellular K+ and increased intracellular Ca+2, abolishing the K+ gradient did not prevent beauvericin-induced apoptosis; in contrast, omission of extracellular Ca+2 inhibited apoptosis by 42%. The ICE family protease inhibitor, Cbz-Val-Ala-Asp-cmk, inhibited apoptosis in a concentration-dependent manner. By Northern blot analysis, cholangiocytes expressed the mRNAs for three members of the ICE protease family: ICE, Ich-1, and CPP-32. Cleavage of a substrate for CPP-32-like protease activity, but not a substrate for ICE and Ich-1, increased following beauvericin treatment. In summary, we have established an in vitro model of apoptosis in cholangiocytes. Our data suggest that beauvericin -induced apoptosis occurs by a Ca+2-dependent, CPP-32 protease -sensitive pathway despite cholangiocyte expression of Bcl-2.

Received 26 April 1996; accepted in final form 28 July 1996.
APS Manuscript Number G162-6.
Article publication pending Am. J. Physiol. (Gastrointest. Liver
Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 21 August 1996