Net h+ and k+ fluxes across the apical surface of rat distal colon. Feldman, George M., and John W. Ickes, Jr. Departments of Medicine and Physiology, McGuire Veterans Affairs Medical Center, Medical College of Virginia / Virginia Commonwealth University, Richmond, VA 23249
APStracts 3:0164G, 1996.
The distal colon absorbs K+ (JK) and secretes H+ (JH) by what is thought to be an H+,K+-ATPase. However, the colonic ATPase differs structurally and functionally from the gastric H+,K+-ATPase. To evaluate the linkage between JH and JK, JH and JK were simultaneously measured with ion specific electrodes in segments of rat distal colon. JH and JK averaged 0.40 0.03 meq h-1 cm-2 and 0.30 0.03 meq h -1 cm-2 (n = 191), but JH and JK did not correlate (r = 0.005, N.S.). The gastric H+,K+ pump inhibitors, SCH28080 (100 mM) and omeprazole (100 mM) as well as a vacuolar H+-ATPase inhibitor, bafilomycin A1 (10 mM), did not affect JH or JK. However, the Na+,K+-ATPase inhibitor, ouabain (1 mM), and N-ethylmaleimide (10 mM) inhibited JK, not JH. Although 1 mM orthovanadate inhibited both JH and JK, at lower concentrations orthovanadate only affected JK. Furthermore, removing K+ from the media did not affect JH. Secondary hyperaldosteronism increased both JH and JK, but ouabain (1 mM) reduced JK, not JH. Cl--free media inhibited voltage insensitive JH and voltage sensitive JK. Medium pH affected JH, but that effect was contrary to the effect that pH had on Rb+ flux (20). These data failed to identify a relationship between JH and JK, and appear to suggest that JH and JK occur by separate pathways.

Received 17 February 1995; accepted in final form 21 August 1996.
APS Manuscript Number G73-5.
Article publication pending Am. J. Physiol. (Gastrointest. Liver
Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 29 August 1996