Programming of hepatic insulin-sensitive enzymes in offspring of rat dams fed an isocaloric protein restricted diet. Desai, Mina, Christopher D. Byrne, Junlong Zhang, Clive J. Petry, Alan Lucas [acute]a, and C. Nicholas Hales . University of Cambridge, Department of Clinical Biochemistry, Addenbrooke's Hospital, Hills Road, Cambridge CB2 2QR, U.K., Dunn Nutritional Laboratory, University of Cambridge and Medical Research Council, Downham's Lane, Milton Road, Cambridge CB4 1XJ, U.K.
APStracts 3:0277G, 1996.
Hepatic enzymes associated with glucose haemostasis were studied in offspring of dams fed either a 20% protein (control) or an isocaloric 8% protein (low protein) diet during pregnancy and lactation. Additionally, offspring were exposed to maternal 8% protein diet only during gestation (recuperated) or lactation (postnatal low protein). The glucokinase activity was decreased (approximately 50%) whereas the phosphoenolpyruvate carboxykinase (PEPCK) activity was increased (about 100%) in the low protein and recuperated offspring compared to the controls (p < 0.001) at 21 days of age. However, the postnatal low protein offspring had enzyme activities comparable to the controls. These changes were still evident in 11 month old offspring weaned onto a normal laboratory chow. Parallel changes were apparent in mRNA levels of glucokinase and PEPCK in the low protein male offspring. Thus the effect of programming metabolism extends not only to protein biochemistry but possibly also to the regulation of gene expression. Furthermore, these changes could not be attributed to glucagon and insulin since their ratio was comparable between the control and low protein groups.

Received 2 July 1996; accepted in final form 6 December 1996.
APS Manuscript Number G266-6.
Article publication pending Am. J. Physiol. (Gastrointest. Liver
Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 31 December 1996