Histamine h2 receptor activation is linked to stimulation of adenylate cyclase and phospholipase c via separate gtp dependent pathways. Wang, L., I. Gantz, and J. Delvalle. Departments of Internal Medicine and Surgery, The University of Michigan Medical Center, Ann Arbor, Michigan 48109
APStracts 3:0085G, 1996.
Previously we demonstrated that a single histamine H2 receptor can couple to both the cAMP and IP3/[Ca2+]i signaling pathways in a stimulatory manner. We undertook the present studies to further characterize the post receptor events involved in H2 receptor dual signaling. Histamine H2 receptor mediated signal transduction was examined in isolated cell membranes prepared from purified canine parietal cells and HEPA cells (rat hepatoma cell line) stably transfected to express the canine H2 histamine receptor cDNA. Histamine dose dependently stimulated both adenylate cyclase (AC, EC50 = 2 x 10-7M) and phospholipase C (PLC, EC50 = 3.1 +/- 0 .5 x 10 -7M) activity in an H2 specific and GTP dependent manner. Cholera toxin pretreatment abolished the stimulatory effect of histamine on PLC activity in isolated membranes without altering binding of the H2 receptor antagonist tiotidine. Anti-Gs[alpha] dose dependently inhibited histamine stimulated AC activity while leaving unaltered the effect of this secretagogue on PLC activity. Although anti -Gq[alpha] inhibited vasopressin stimulated PLC activity in HEPA cells and carbachol stimulated PLC in parietal cells, this antibody did not alter the action of histamine on PLC in the same membrane preparations. Antibody against the amino and carboxyl terminus of the common [beta] subunit of heterotrimeric G-proteins did not inhibit histamine stimulated PLC activity. Our studies demonstrate for the first time that activation of the H2 receptor leads to stimulation of both AC and PLC via separate GTP dependent mechanisms.

Received 22 July 1995; accepted in final form 8 April 1996.
APS Manuscript Number G320-5.
Article publication pending Am. J. Physiol. (Gastrointest. Liver
Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 1 May 96