Physiological variations in ovine cerebrovascular calcium sensitivity. Akopov, Sergey E., Lubo Zhang, and William J. Pearce. Departments of Physiology, Pharmacology, and Biochemistry, Center for Perinatal Biology, Loma Linda University School of Medicine, Loma Linda, California, 92350
APStracts 3:0542H, 1996.
Cerebrovascular reactivity to biogenic amines varies in relation to both artery diameter and age. The present study examines the hypothesis that these patterns of reactivity are secondary to corresponding variations in the Ca2+ sensitivity of the contractile proteins. To test this hypothesis, we permeabilized segments of common carotid (COM), basilar (BAS), main branch middle cerebral (MCA), and second branch middle cerebral (MCA-B) arteries from non -pregnant adult and near-term fetal sheep using -escin. Permeabilization methods were carefully validated and adjusted for each artery type. Baseline myofilament Ca2+ sensitivity in both adults and fetuses increased significantly from COM to MCA-B, and was generally higher in fetuses than adults. Serotonin dose-dependently increased Ca2+ sensitivity via a G-protein dependent mechanism in all arteries. The magnitudes of this effect did not vary among artery types but were significantly greater in fetal than adult arteries. This effect of 5HT was mimicked by GTPgS, a nonhydrolyzable analog of GTP, and its effects were also much greater in fetal than adult arteries. We conclude that patterns of cerebrovascular reactivity to biogenic amines are determined, at least in part, by underlying variations in baseline myofilament Ca2+ sensitivity and/or its alteration by G-protein dependent mechanisms.

Received 27 September 1996; accepted in final form 6 December
1996.
APS Manuscript Number H874-6.
Article publication pending Am. J. Physiol. (Heart Circ. Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 31 December 1996