Inward rectifier k+ currents in smooth muscle cells from rat coronary arteries: block by mg2+, ca2+, and ba2+ ions. Robertson, Blair E., Adrian D. Bonev, and Mark T. Nelson. Department of Pharmacology, The University of Vermont, 55A South Park Drive, Colchester, Vermont 05446, USA
APStracts 3:0014H, 1996.
Inward rectifier potassium channels have been implicated in the control of membrane potential and external potassium-induced dilations of small coronary arteries. To identify and characterize inward rectifier K+ currents in coronary artery smooth muscle, whole cell K+ currents in smooth muscle cells enzymatically isolated from rat coronary (septal) arteries (diameters, 100-150 microns) were measured in the conventional and perforated configurations of the patch clamp technique. Barium-sensitive, whole cell K+ current -voltage relationships exhibited inward rectification. Blockers of calcium-activated K+ channels (1 mM TEA+), ATP-sensitive K+ channels (10 [mu]M glibenclamide) and voltage-dependent K+ channels (1 mM 4 -aminopyridine) in smooth muscle were without affect on inward rectifier K+ currents. The non-selective K+ channel inhibitor, phencyclidine (100 [mu]M) reduced inward rectifier K+ currents by about 50%. External barium reduced inward currents, with membrane potential hyperpolarization increasing inhibition. The half -inhibition constant for barium was 2.1 [mu]M at -60 mV, decreasing e -fold for a 25 mV hyperpolarization. External cesium also blocked inward rectifier K+ currents, with the half-inhibition constant for cesium of 2.9 mM at -60 mV. External calcium and magnesium reduced inward rectifier K+ currents. At -60 mV, calcium and magnesium (1 mM) reduced inward currents by 33% and 21%, respectively. Inward rectification was not affected by dialysis of the cell's interior with a nominally calcium- and magnesium-free solution. These findings indicate that inward rectifier K+ channels exist in coronary artery smooth muscle and that barium may be a useful probe for the functional role of inward rectifier K+ channels in coronary arteries.

Received 6 April 1995; accepted in final form 12 December 1995.
APS Manuscript Number H336-5.
Article publication pending Am. J. Physiol. (Heart Circ. Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 22 January 96