Adisrupter of actin microfilaments impairs sulfonylurea-inhibitory gating of cardiac katp channels. Brady, Peter A., Alexey E. Alekseev, Luba A. Aleksandrova, Luis A. Gomez, and Andre Terzic. Division of Cardiovascular Diseases, Department of Medicine and Pharmacology, Mayo Clinic, Mayo Foundation, Rochester, MN 55905
APStracts 3:0413H, 1996.
The efficacy with which sulfonylureas inhibit cardiac ATP-sensitive K+ (KATP) channels is reduced during metabolic compromise and cellular contracture. Disruption of the actin-microfilament network, which occurs under similar conditions, reduces the sensitivity of the channel towards intracellular ATP. To investigate whether a disrupter of actin-microfilaments could also affect the responsiveness of the KATP channel to sulfonylureas, single channel currents were measured in the inside-out configuration of excised patches from guinea-pig ventricular myocytes. Treatment of the internal side of patches with DNase I (100 [mu]g/ml), which forms complexes with G-actin and prevents actin-filament formation, antagonized sulfonylurea-induced inhibition of KATP channels which was coupled with a loss of sensitivity to ATP. The apparent Kd and Hill coefficient for the inhibitory effect of glyburide, a prototype sulfonylurea, on KATP channel opening were respectively 0.13 [mu]M and 0.95 prior to and 2.7 [mu]M and 0.98, following DNase-treatment. DNase did not alter intraburst kinetic properties of the channel. When denatured or co -incubated with purified actin (200 [mu]g/ml), DNase I no longer decreased glyburide-induced channel inhibition. This suggests that sulfonylurea-inhibitory gating of cardiac KATP channels may also be regulated through a mechanism involving sub-sarcolemmal actin -microfilament networks.

Received 7 August 1996; accepted in final form 13 September 1996.
APS Manuscript Number H717-6.
Article publication pending Am. J. Physiol. (Heart Circ. Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 5 November 1996