Endothelial cytoskeleton and lung capillary integrity - ultrastructural changes in response to botulinum c2 toxin. Ermert, L., H. R. Duncker, H. Br[umlaut]uckner, F. Grimminger, T. Hansen, R. R[diaeresis]ossi, K. Aktories, and W. Seeger. Department of Internal Medicine, Justus-Liebig-University, Giessen, 35385 Giessen, and the Institute of Anatomy and Cellbiology, Justus -Liebig-University, Giessen, 35385 Giessen, and the Institute of Pharmacology and Toxicology, University of the Saarland, 66421 Homburg, Germany
APStracts 3:0474A, 1996.
The role of the actin microfilament system in endothelial cells for the structural integrity of the pulmonary gas exchange area was probed by use of Clostridium botulinum C2 toxin. This agent causes selective loss of non-muscle F-actin. Buffer-perfused rabbit lungs were used, in which the vascular pressures varied within physiological ranges. Group A (n=3) included control lungs, buffer -perfused for 180 min. Group B lungs (n=3) were poisoned with low-dose C2 toxin (0.3/0.6 ng/ml C2I/II in the buffer fluid; n=3), and experiments were terminated after a total weight gain due to edema accumulation of 7.5 g (122 +/- 16 min). Lungs of groups C and D were exposed to high-dose C2 toxin (10/20 ng/ml C2I/II; n=3 each), and perfusion was stopped after a total weight gain of 1 g (16 +/- 3 min; C) and 7.5 g (38 +/- 6 min; D). Perfusion fixation for electron -microscopic evaluation was performed immediately after cessation of buffer circulation. Group A lungs did not gain weight and displayed normal lung architecture. In low-dose C2 toxin-treated lungs (B), extensive attenuations, undulations and protrusions of the endothelial layer were noted, suggestive of "remodeling" and "flowing" of the cell membranes. These were accompanied by few disruptions of the endothelial layer, edema formation in the vicinity of these lesions as well as vesiculation and bleb formation of endothelial cells. Lungs that were exposed for short-term periods to the high toxin doses (C, D) again displayed marked attenuations of the endothelial layer, including both the thin and the thick side of the septal capillary walls, in addition to large endothelial cell disruptions, particularly in group D. These lesions did not include interendothelial junctions, although they were often located adjacent to them in any of the C2 toxin-treated lungs. Significant ultrastructural changes of interstitial cells and epithelial type-I cells were not seen, interestingly type-II epithelial cells displayed fusion of lamellar bodies. Collectively, these data suggest that the actin microfilament system is instrumental in supporting endothelial cell membrane configuration and integrity and maintains the intimal barrier function of the lung microvasculature.

Received 13 February 1996; accepted in final form 4 September
1996.
APS Manuscript Number A150-6.
Article publication pending Journal of Applied Physiology.
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 5 November 1996