Instantaneous force-velocity-length relationship in diaphragmatic sarcomere. Coirault, Catherine, Denis Chemla, Jean-Claude Pourny, Francine Lambert, Yves Lecarpentier. INSERM 451-Laboratoire d'Optique Appliqu_e-Ecole Polytechnique, 91125 Palaiseau Cedex and Service d'Explorations Fonctionnelles, Centre Hospitalier et Universitaire de Bic_tre, 94275 Le Kremlin -Bic_tre, France
APStracts 3:0478A, 1996.
The simultaneous analysis of muscle force, length, velocity and time has been shown to precisely characterize the mechanical performance of isolated striated muscle. We tested the hypothesis that the three -dimensional force-velocity-length relationship, reflects mechanical properties of sarcomeres. In hamster diaphragm strips, instantaneous sarcomere length (SL) and muscle length were simultaneously measured during afterloaded twitches. SL was measured by means of laser diffraction. We also studied the influence of initial sarcomere length, abrupt changes in total load, and 2 x 10-7 M dantrolene. Baseline resting SL at the apex of the length-active tension curve was 2.2 +/- 0.1 [mu]m, while SL at peak shortening was 1.6 +/- 0.1 [mu]m in the preloaded twitch and 2.1 +/- 0.1 [mu]m in the "isometric" twitch. Over the whole load continuum and at any given level of isotonic load, there was a unique relationship between instantaneous sarcomere velocity (SV) and instantaneous SL. Part of this relationship was time- and initial SL-independent and was markedly down-shifted after dantrolene. When five different muscle regions were considered, there was no significant variations of SL and sarcomere kinetics along the muscle. These results indicate that the time- and initial length-independent part of the instantaneous force-velocity-length relationship previously described in muscle strips reflects intrinsic sarcomere mechanical properties.

Received 9 May 1996; accepted in final form 1 October 1996.
APS Manuscript Number A444-6.
Article publication pending Journal of Applied Physiology.
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 5 November 1996