The cns site of glucocorticoid negative feedback during lipopolysaccharide and psychological stress-induced fevers. Morrow, Lee E., Jennifer L. McClellan, John J. Klir, Matthew J. Kluger. Department of Physiology, The University of Michigan Medical School, Ann Arbor, MI 48109, College of Medicine, The University of Iowa, Iowa City, IA 52242, Department of Physiology, Ohio State University College of Medicine, Columbus, OH 43210, Institute for Basic and Applied Medical Research, The Lovelace Institutes, 2425 Ridgecrest Dr. SE, Albequerque, NM 87108, Phone: 505 262-7531, FAX: 505 262-7043
APStracts 3:0153R, 1996.
The CNS Site of Glucocorticoid Negative Feedback During Lipopolysaccharide and Psychological Stress-induced Fevers. Glucocorticoids exert negative feedback in the anterior hypothalamus (AH) during lipopolysaccharide (LPS)-induced fevers, but the central location of their negative feedback during psychological stress -induced fever has not been determined. To confirm that glucocorticoid modulation of LPS fever occurs in the AH, adrenalectomized animals were injected intrahypothalamically (ih) with either 0.25 ng of corticosterone or vehicle followed by 50 [mu]g/kg LPS intraperitoneally (ip). Animals pretreated with corticosterone developed significantly smaller fevers (P = 0.007) than animals given vehicle. To determine if glucocorticoid modulation during psychological stress-induced fever may occur in the hippocampus, the fornix was transected to block hippocampal communication with the AH. This resulted in significantly larger psychological stress-induced fevers (P = 0.02) compared to sham operated animals. There were no differences between these groups for LPS-induced fevers (P = 0.92). To determine where in the hippocampus glucocorticoids might exert their negative feedback during psychological stress, rats were microinjected with either 1 ng RU38486 (a type II glucocorticoid receptor antagonist) or vehicle into the dentate gyrus prior to exposure to the open-field. There were no differences between the psychological stress-induced fevers of the RU38486 and vehicle -injected groups, supporting the hypothesis that these fevers are modulated elsewhere in the hippocampus. Our data support the hypothesis that glucocorticoids modulate LPS-induced fever in the anterior hypothalamus and do not involve the hippocampus, and that psychological stress-induced fevers are modulated by neural connections between the hippocampus and the hypothalamus. The precise sites of action of glucocorticoid negative feedback on stress-induced fevers in the hippocampus (or other brain regions) is not yet known.

Received 1 June 1995; accepted in final form 8 April 1996.
APS Manuscript Number R330-5.
Article publication pending Am. J. Physiol. (Regulatory Integrative
Comp. Physiology).
ISSN 1080-4757 Copyright 1996 The American Physiological Society.
Published in APStracts on 1 May 96